QTLdb: entrichment tests

Release 59
(Apr 20, 2026)

Whole genome analysis for QTL/association enrichment

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Version: Enrich S: beta v0.8

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Data:

Number of milk composition - other traits:	29
Number of QTL / associations found:	2,720
Number of chromosomes where QTL / associations are found:	30

Chi-squared (χ²) test: are milk composition - other traits over-represented on some chromosomes?

Chromosomes	Total χ²	df	p-values	FDR ^*	Size of χ²
Chromosome X	2402.45106	29	9e-41	1.227273e-40
Chromosome 1	2439.55395	29	9e-41	1.227273e-40
Chromosome 2	22011.88856	29	9e-41	1.227273e-40
Chromosome 3	164.33343	29	7.535731e-21	9.419664e-21
Chromosome 4	1508.14210	29	1.001237e-299	3.003711e-298
Chromosome 5	12.82960	29	0.9958835	9.983293e-01
Chromosome 6	73489.55395	29	9e-41	1.227273e-40
Chromosome 7	164.33343	29	7.535731e-21	9.419664e-21
Chromosome 8	430.02447	29	5.945344e-73	1.274002e-72
Chromosome 9	362.53567	29	2.714242e-59	5.428484e-59
Chromosome 10	111.45106	29	1.331564e-11	1.536420e-11
Chromosome 11	159.53567	29	5.593902e-20	6.712682e-20
Chromosome 12	78.50619	29	1.912381e-06	2.048980e-06
Chromosome 13	99.82960	29	1.041720e-09	1.157467e-09
Chromosome 14	21016.50619	29	9e-41	1.227273e-40
Chromosome 15	821.09817	29	4.287883e-154	1.429294e-153
Chromosome 16	887.20106	29	5.38336e-168	2.307154e-167
Chromosome 17	696.56869	29	5.149778e-128	1.404485e-127
Chromosome 18	1421.56869	29	2.841781e-281	2.841781e-280
Chromosome 19	853.82960	29	5.66654e-161	2.124953e-160
Chromosome 20	667.03567	29	7.439992e-122	1.859998e-121
Chromosome 21	1296.50619	29	1.178875e-254	8.841563e-254
Chromosome 22	1642.80012	29	9e-41	1.227273e-40
Chromosome 23	1642.80012	29	9e-41	1.227273e-40
Chromosome 24	638.14210	29	7.707772e-116	1.778717e-115
Chromosome 25	1.74145	29	0.998329325823115	9.983293e-01
Chromosome 26	1464.53567	29	1.985029e-290	2.977543e-289
Chromosome 27	955.86262	29	1.809059e-182	9.045295e-182
Chromosome 28	757.55395	29	9.113352e-141	2.734006e-140
Chromosome 29	1027.08343	29	1.63124e-197	9.787440e-197

Chi-squared (χ²) test: Which of the 29 milk composition - other traits are over-represented in the QTLdb

Traits	Total χ²	df	p-values	FDR ^*	Size of χ²
Milk acetone content	69.17178	28	2.427284e-05	3.704802e-05
Milk acidity/alkalinity	65.89283	28	6.831806e-05	9.277507e-05
Milk beta-hydroxybutyrate content	97.53224	28	1.267506e-09	2.162216e-09
Milk calcium content	300.93236	28	1.602296e-47	5.162954e-47
Milk cholesterol content	128.82374	28	6.989151e-15	1.447753e-14
Milk citrate content	63.97418	28	0.0001235286	1.557535e-04
Milk copper content	66.52844	28	5.602255e-05	8.123270e-05
Milk energy yield	65.79723	28	7.038109e-05	9.277507e-05
Milk glycerophosphocholine content	906.3474	28	8.688306e-173	6.299022e-172
Milk glycerophosphocholine to phosphocholine ratio	732.95785	28	2.478691e-136	1.437641e-135
Milk iodine content	95.4899	28	2.692249e-09	4.337512e-09
Milk iron content	377.46281	28	7.202784e-63	2.611009e-62
Milk lactose content	194.80242	28	6.571928e-27	1.905859e-26
Milk lactose yield	60.92273	28	0.0003100617	3.596716e-04
Milk magnesium content	151.46848	28	6.69829e-19	1.494234e-18
Milk manganese content	191.28572	28	3.017728e-26	7.955828e-26
Milk mid-infrared spectra	115.93407	28	1.148046e-12	2.219556e-12
Milk phosphocholine level	62.34898	28	0.0002023608	2.445193e-04
Milk phosphorus content	1264.79364	28	9.575076e-249	2.776772e-247
Milk potassium content	1185.8459	28	5.769563e-232	5.577244e-231
Milk riboflavin content	553.5835	28	5.826293e-99	2.816042e-98
Milk selenium content	426.174	28	9.154832e-73	3.792716e-72
Milk sodium content	55.58708	28	0.001443959	1.610570e-03
Milk solids percentage	111.09719	28	7.496771e-12	1.358790e-11
Milk solids yield	27.98877	28	0.4650428	4.994904e-01
Milk vitamin B-12 content	163.02306	28	5.319285e-21	1.285494e-20
Milk zinc content	1227.28987	28	9.0223e-241	1.308233e-239
Non-fat milk solids	26.15617	28	0.5644606	5.846199e-01
Non-fat milk solids percentage	6.83863	28	0.9999855	9.999855e-01

Correlations found between some of these traits for your reference

No correlation data found on these traits

Overall Test

Data		Chi'Square Test			Fisher's Exact Test
Number of chrom.:	30	χ²	=	139225.802460
Number of traits:	29	df	=	812
Number of QTLs:	2,720	p-value	=	0

FOOT NOTE: * : FDR is short for "false discovery rate", representing the expected proportion of type I errors. A type I error is where you incorrectly reject the null hypothesis, i.e. you get a false positive. It's statistical definition is FDR = E(V/R | R > 0) P(R > 0), where V = Number of Type I errors (false positives); R = Number of rejected hypotheses. Benjamini–Hochberg procedure is a practical way to estimate FDR.