- Tian R, Cullen NG, Morris CA, Fisher PJ, Pitchford WS, Bottema CD (2012). Major effect of retinal short-chain dehydrogenase reductase (RDHE2) on bovine fat colour. Mammalian genome,23 (5-6): 378-86. [see all QTL
 by this paper]
- Tian R, Pitchford WS, Morris CA, Cullen NG, Bottema CD (2010). Genetic variation in the beta, beta-carotene-9', 10'-dioxygenase gene and association with fat colour in bovine adipose tissue and milk. Animal genetics,41(3): 253-9. [see all QTL by this paper]
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